is a German zoologist and molecular biologist. He earned his doctorate in Freiburg i. Br. Since 2003 he has been a group leader at the European Molecular Biology Lab in Heidelberg. His main area of research is the evolution of animal body plans; his work on the molecular evolution of cell types has pioneered the growing field of cell type evolution in animals. In 2011 he received the Kovalevsky Medal from the St. Petersburg Society of Naturalists.